- John Leguizamo Character (Sid (voice))
- Ray Romano Character (Manny (voice))
- Chris Wedge Character (Scrat (voice))
- Denis Leary Character (Diego (voice))
- Seann William Scott Character (Crash (voice))
- Jennifer Lopez Character (Shira (voice))
- Queen Latifah Character (Ellie (voice))
- Peter Dinklage Character (Captain Gutt (voice))
- Nick Frost Character (Flynn (voice))
- Josh Gad Character (Louis (voice))
- Nicki Minaj Character (Steffie (voice))
- Heather Morris Character (Katie (voice))
- Wanda Sykes Character (Granny (voice))
- Alan Tudyk Character (Milton / Hunky Siren (voice))
- Rebel Wilson Character (Raz (voice))
- Aziz Ansari Character (Squint (voice))
- Joy Behar Character (Eunice (voice))
- Christopher Campbell Character (Creature Siren (voice))
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