- Ben Stiller Character (Alex (voice))
- Jada Pinkett Smith Character (Gloria (voice))
- David Schwimmer Character (Melman (voice))
- Chris Rock Character (Marty / Additional Zebras (voice))
- Cedric the Entertainer Character (Maurice (voice))
- Andy Richter Character (Mort (voice))
- Bernie Mac Character (Zuba (voice))
- Alec Baldwin Character (Makunga (voice))
- Sacha Baron Cohen Character (Julien (voice))
- Sherri Shepherd Character (Mom (voice))
- Will.i.am Character (Moto Moto (voice))
- Elisa Gabrielli Character (Nana (voice))
- Tom McGrath Character (Skipper / Lemur (voice))
- Chris Miller Character (Kowalski (voice))
- Christopher Knights Character (Private (voice))
- Conrad Vernon Character (Mason (voice))
- Quinn Dempsey Stiller Character (Baby Alex (voice))
- Declan Swift Character (Baby Alex (voice))
- Fred Tatasciore Character (Teetsi / Poacher #1 / Elephant (voice))
- Eric Darnell Character (Joe the Witch Doctor / Poacher #2 (voice))
- Willow Smith Character (Baby Glorida (voice))
- Thomas Stanley Character (Baby Marty (voice))
- Zachary Gordon Character (Baby Melman (voice))
- Meredith Vieira Character (Newscaster (voice))
- Lesley Stahl Character (Newscaster (voice))
- Al Roker Character (Newscaster (voice))
- David Soren Character (Lemur (voice))
- Phil LaMarr Character (Guide (voice))
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